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Broken Tusk, now recovered due to the intervention of a human doctor, sides with Machiko, and together with the cargo ship's crew they arrange for the transporter's massive orbiter to crash into Ryushi and destroy the colony and the Aliens. The Predator assault continues to the settler colony itself, and the surviving settlers find themselves pitched between the Aliens and Predators. Meanwhile, the infected Rhynth have been loaded aboard a cargo transporter and, with a queen among their number, an Alien colony quickly takes hold. However, they soon encounter the settlers and, after Broken Tusk is incapacitated, change their plans to hunt them instead. Led by a Predator elder, 'Broken Tusk', the Predators arrive expecting to encounter Aliens. On reaching Ryushi, the eggs hatch and infect Rhynths. Confounding the Predator's safeguards, this queen manages to slip an egg containing the seed of another queen into the shipment. On board the Predator ship, the prey are prepared: an Alien queen lays eggs for delivery to Ryushi. Unbeknownst to the colonists, Ryushi is a traditional hunting ground of the Predators, and they are returning for their initiation rites. The settlers on Ryushi raise cattle-like quadrupedal ungulates called rhynth for export to other solar systems, and at the time of the story are in the process of assembling a shipment of the native livestock.

The first Aliens versus Predator centers on Ryushi, a recently colonized planet, and Machiko Noguchi, the Chigusa Corporation's administrator there.

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